The piece shown is a Yoruba beaded crown. These were worn by Yoruba kings (sort of like Greece the Yoruba had a number of kings ruling city states….but I studied all that a long time ago so I have no more details for you).
The crowns are covered with two symbols from Yoruba culture—birds and mysterious faces. The faces are a representation of the creator god, Oduduwa and the birds, called Okin, represent women. Apparently the king, by wearing the crown, acknowledges that he cannot rule without the support of god…and women. This particular piece sells for $1995 (on sale from August 1-5).
For more info head HERE.